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한국 자포니카 품종에서 벼퉁그로 바이러스병 저항성 탐색

이종희1,*, 조준현1, 이지윤1, 강주원1, 신동진1, 조수민1, 최일룡2, 송유천1, 박동수1, 고종민1

Identification of Resistance to Rice Tungro Virus Disease in Korean Japonica Rice Cultivars

Jong Hee Lee1,*, Jun-Hyeon Cho1, Ji-Yoon Lee1, Ju-Won Kang1, Dongjin Shin1, Sumin Jo1, Il-Ryong Choi2, You-Chun Song1, Dong-Soo Park1, Jong-Min Ko1
Korean Journal of Breeding Science 2019;51(2):86-90.
Published online: June 1, 2019

농촌진흥청 국립식량과학원

국제미작연구소

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*(E-mail: ccriljh@korea.kr, Tel: +82-55-350-1168, Fax: +82-55-352-3059)
• Received: April 29, 2019   • Revised: April 30, 2019   • Accepted: May 16, 2019

Copyright: © 2019 by the Korean Society of Breeding Science

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  • Rice tungro disease is a serious threat to rice production in South and Southeast Asian countries. Rice production in these countries has been intensified, often through continuous cultivation of rice. We conducted a screen of the resistance to tungro virus disease in Korean japonica rice. Dongjin, Hwaseong, and Sangju were varieties resistant to Rice tungro spherical virus (RTSV) but susceptible to Rice tungro bacilliform virus (RTBV). RTSV-resistant variety Sangju was crossed with RTSV-susceptible variety Unkwang to conduct a genetic analysis for RTSV resistance originating from Sangju. Evaluation of 201 F2 plants from the cross between Unkwang and Sangju showed that there were 85 plants showing resistance, and that 116 plants were susceptible, indicating a ratio fitting a 7:9 ratio ratio (χ2 = 0.54; p = 0.46). Thus, the resistance to RTSV in Sangju might be controlled by two recessive genes. This result indicates that Sangju might serve as a useful genetic source for diversification of RTSV resistance and stabilization of resistance through gene pyramiding.
  • Keywords: rice; RTSV; Sangju; tungro virus disease
지구 온난화에 따른 한반도의 최근 30년간 평균온도는 1.7°C 상승하였으며, 이로 인한 벼에서 병해충 발생량은 점점 증가하고 있는 실정이다. 특히, 바이러스병인 줄무늬잎마름병은 1960년대 낙동강 주변에서 일부 남부지역에서 발생되었으나, 2000년 이후는 발생지역이 북상하여 전국으로 확대되었고, 2007년에는 중국에서 바이러스 보독충이 비래하여 큰 피해를 입힌 바 있다(Lee et al. 2008). 또한 최근 기온상승에 따른 한반도와 주변지역의 아열대화 현상이 가속화 되고 있으며, 바이러스병 저항성 육종에 있어도 이에 대응한 선제적인 대응책의 마련이 필요한 실정이다.
동남아시아 아열대 지역에서 가장 심각한 바이러스병은 벼퉁그로 바이러스병(rice tungro disease, RTD)이며, 온대지역인 일본 남부지역에 Waikai 바이러스란 이름으로 발생이 보고된 바 있다. 이병은 Rice tungro bacilliform virus (RTBV)와 Rice tungro spherical virus (RTSV)의 상호작용에 의해 두점끝동매미충으로 전염된다. RTBV는 벼의 엽색이 노란색 또는 연황색으로 변색되면서 분얼수가 작아지고 위축되는 병징에 관여하고 있지만, 이병 단독으로 병징이 나타나지 않았고, RTSV와 RTBV가 함께 감염되어야 병징을 나타낸다. 그리고 RTSV 단독으로 감염된 경우는 뚜렷한 병징은 나타나지 않으며, 생육저해 현상이 크지 않은 편이다. 반면, RTSV는 두점끝동매미충에 의해 바이러스를 식물체로 쉽게 전염시키지만, RTBV는 RTSV를 흡즙한 두점끝동매미충에 의해서 전염되고, RTSV를 보유하지 않을 경우 전염시키지 못한다(Hibino et al. 1979). 즉 RTSV에 저항성인 품종은 두점끝동매미충이 RTBV에 보독되는 것을 사전에 차단함으로써 벼퉁그로 바이러스병 발생을 억제할 수 있다.
RTSV에 저항성인 품종은 Utri Merah82, Ultri Rajapan, Pankari 203, TKM6 등이 보고 되었으며, 국제미작연구소에서는 대부분 TKM6에서 유래된 저항성 품종이 이용되었다고 하였다. 또한 야생벼에서 유래된 Matatag 9은 야생벼 O. rufipogon에서 저항성이 유래되었다고 알려져 있으며, 최근 RTSV에 진성저항성이 아닌 매개충인 두점끝동매미충에 저항성을 나타내는 것으로 밝혀졌다. Imbe et al. (1993)은 Waikai 바이러스에 저항성인 17 자포니카 품종에 대해 RTSV 저항성 검정을 수행한 결과, 모두 저항성 반응을 나타내었으며, 이는 일본 재래도 Takenari에서 유래되었다고 하였다.
국내에서 퉁그로바이러스에 대한 발생이 보고되지 않았지만, 한반도의 온도 상승과 더불어 발병 위험은 점점 증가되고 있는 실정이다. 따라서, 본 연구에서는 국제미작연구소와 공동으로 국내육성품종에 대해 퉁그로바이러스에 대한 저항성 탐색과 유전분석을 수행하고, 향후 이병에 대한 저항성 육종기반을 구축하고자 연구를 수행하였다.
한국 자포니카 품종에 대한 퉁그로바이러스병 저항성 검정을 수행하기 위해 국제미작연구소 GUVA project에서 교배친으로 활용하고 있는 자포니카형 89품종을 이용하였다. 그리고, 선발된 저항성 품종의 유전분석을 위해 감수성 품종인 운광벼와 저항성 품종인 상주벼가 교배된 F1 식물체 및 F2 집단을 이용하여 바이러스 생물검정을 수행하였으며, 카이스퀘어검정(Chi-square test)을 통한 분리비 적합도 검정을 수행하였다.
퉁그로바이러스 저항성 검정은 감수성 품종인 TN1에서 유지된 RTSV 균주 A와 RTBV common strain를 접종원으로 사용하였다. 식물체에 바이러스 접종은 Cabauatan et al. (1995)의 방법에 따라 두점끝동매미충(GLH)을 RTSV와 RTBV가 이병된 식물체에 3일간의 흡즙하여 보독충을 만들었다. 묘령이 6-10일된 식물체에 주당 바이러스 보독충 3마리를 24시간동안 접종하였다. 접종된 식물체는 국제미작연구소 온실에서 유지하였으며, 접종 후 14일째와 21일째 잎에서 즙액을 채취하였으며, ELISA 판독기(BioTek Instruments, Winooski, USA)로 흡광도 405 nm에서 측정하여 저항성과 감수성을 결정하였다. 10배 희석 된 추출물이 0.1보다 큰 흡광도 값을 나타내는 식물체는 바이러스에 감염된 것으로 간주되었다. 시험 품종당 최소 12개 이상 측정하였으며, 감염률이 20% 미만인 품종은 퉁그로바이러스에 저항성인 것으로 판단하였다(Cabauatan et al. 1995).
한국 자포니카 품종을 검정한 결과, RTSV에 이병율이 20% 이하인 3품종이 있었으며, 이병율이 21%~60% 사이는 56품종, 61% 이상이 30품종이 있었다. 반면 RTBV에 대해서는 이병율이 80% 이상으로 모두 감수성 반응을 나타내었다(Table 1).
Table 1
Infection ratio of rice tungro spherical virus (RTSV) and rice tungro bacilliform virus (RTBV) in Korean Japonica varieties.
Table 1
Entry No. Varieties RTBV RTSV Entry No. Varieties RTBV RTSV
1 Aranghyangchalbyeo 73.7 89.5 46 Jongnambyeo 100.0 37.8
2 Ansanbyeo 100.0 45.5 47 Juanbyeo 100.0 36.4
3 Anseongbyeo 100.0 83.3 48 Junganbyeo 100.0 27.5
4 Bongkwangbyeo 100.0 58.8 49 Junghwabyeo 100.0 62.5
5 Cheonmabyeo 100.0 27.8 50 Jungsangbyeo 100.0 65.8
6 Chucheongbyeo 100.0 73.0 51 Kwanganbyeo 100.0 63.2
7 Daeanbyeo 100.0 75.0 52 Mananbyeo 100.0 26.3
8 Daechangbyeo 100.0 38.5 53 Manchubyeo 100.0 62.5
9 Daejinbyeo 100.0 29.6 54 Mangeumbyeo 100.0 45.5
10 Daepyungbyeo 95.0 85.0 55 Manhobyeo 100.0 52.2
11 Daeribbyeo 1 100.0 41.2 56 Manmibyeo 100.0 25.9
12 Daesanbyeo 100.0 24.3 57 Manpungbyeo 100.0 41.0
13 Daeyabyeo 100.0 69.2 58 Mihyangbyeo 100.0 61.1
14 Dongjin1 100.0 55.0 59 Naepungbyeo 100.0 64.7
15 Dongjinbyeo 100.0 8.6 60 Nampyeongbyeo 100.0 54.3
16 Dongjinchalbyeo 100.0 51.4 61 Namweonbyeo 87.1 45.2
17 Dunnaebyeo 100.0 26.7 62 Nonganbyeo 100.0 34.3
18 Gancheokbyeo 100.0 42.5 63 Nongbaek 100.0 47.2
19 Geumanbyeo 100.0 54.5 64 Obongbyeo 100.0 26.7
20 Geumnambyeo 100.0 34.3 65 Odaebyeo 100.0 61.1
21 Geumobyeo 87.2 46.2 66 Palgongbyeo 100.0 52.9
22 Geumobyeo 100.0 76.2 67 Saegyehwabyeo 84.6 61.5
23 Goamy 100.0 62.5 68 Sambaekbyeo 100.0 37.0
24 Gurubyeo 100.0 78.8 69 Samcheonbyeo 100.0 28.2
25 Haepyeongbyeo 100.0 33.3 70 Sangjubyeo 94.1 0.0
26 Heughyangbyeo 100.0 47.2 71 Sangmibyeo 100.0 64.3
27 Heugjinjubyeo 97.1 37.1 72 Sangnambadtbyeo 89.2 48.6
28 Heugnambyeo 100.0 55.0 73 Sanjuchalbyeo 100.0 44.1
29 Hoanbyeo 100.0 25.0 74 Seoanbyeo 100.0 28.6
30 Hwaanbyeo 100.0 44.4 75 Seojinbyeo 87.9 60.6
31 Hwabongbyeo 100.0 63.2 76 Seolgengbyeo 100.0 55.3
32 Hwadongybeo 100.0 21.1 77 Seolhyangchalbyeo 100.0 32.4
33 Hwajinbyeo 100.0 63.6 78 Seomjinbyeo 100.0 35.0
34 Hwajungbyeo 100.0 72.2 79 Sinseonchalbyeo 100.0 30.8
35 Hwamyeongbyeo 100.0 62.9 80 Sobaekbyeo 100.0 53.6
36 Hwanambyeo 100.0 91.7 81 Sobibyeo 100.0 80.0
37 Hwasambyeo 100.0 61.5 82 Sujinbyeo 100.0 41.7
38 Hwaseongbyeo 92.1 7.9 83 Taeseongbyeo 94.3 62.9
39 Ilmibyeo 92.5 25.0 84 Tamjinbyeo 100.0 65.2
40 Ilpumbyeo 100.0 48.6 85 Umbongbyeo 100.0 45.7
41 Janganbyeo 100.0 21.6 86 Undoobyeo 73.7 78.9
42 Jinbongbyeo 100.0 58.3 87 Unjangbyeo 100.0 35.3
43 Jinbuolbyeo 100.0 37.8 88 Weonhwangbyeo 100.0 38.5
44 Jinheung 100.0 82.4 89 Yeonganbyeo 100.0 35.7
45 Jinjubyeo 100.0 68.0
국내 육성 자포니카 품종에서 RTSV 이병율이 20% 이하로 저항성을 보인 품종은 상주벼, 화성벼, 동진벼 였으며, 이들의 바이러스 이병율을 Table 2에 나타내었다. 상주벼는 RTSV와 RTBV가 복합 감염되거나, RTSV 단독으로 감염된 개체수는 없었으며, 다른 품종보다 강한 저항성 반응을 보였다. 화성벼와 동진벼는 RTSV에 단독으로 감염된 개체는 없었으나, RTSV와 RTBV가 복합적으로 감염된 개체수가 있었고, 이들 각각 7.9%와 8.9%이였다(Table 2).
Table 2
The infection ratio of moderate resistant varieties against to rice tungro spherical virus (RTSV).
Table 2
Varieties Infection ratio (%)
RTBV+RTSV RTBVz RTSV
Sangjubyeo 0.0 94.1 0.0
Hwaseongbyeo 7.9 84.2 0.0
Dongjinbyeo 8.6 85.7 0.0

zRice tungro bacilliform virus

국제미작연구소에서 RTSV저항성 유전자원으로 활용된 품종은 TKM6, Pnakahari 203 및 Utri Merah 82 등이 활용되었으며, 이들 인디카 배경의 품종은 RTSV에 매우 강한 저항성 반응을 나타내었다(Imbe et al. 1993, Shim et al. 2015). 또한, 벼 퉁그로 바이러스병과 유사한 Waikai 바이러스병에 저항성인 품종에 대해 RTSV 저항성 검정을 수행한 결과 Aichi Asahi, Kinmaze, Nishikaze, Saikai 139가 저항성 반응을 나타내었고, 이들은 일본의 재래품종인 Takenari에서 유래되었다고 하였다. 본 연구에서 저항성으로 탐색된 동진벼의 유전적 계보를 추적한 결과 일본 품종의 저항성원으로 사용된 Kinmaze가 존재하였다. 따라서 동진벼는 일본의 재래품종인 Takenari에서 유래되었을 것으로 판단된다. Lee et al. (2011)은 Utri Merah 82유래의 RTSV 저항성 유전자 tsv1를 분리 동정하였으며, translation initiation factor (eIF4G) 유전자에 저항성이 결정된다고 보고하였다. 또한 이 유전자 영역의 SNP 분석을 한 결과 TKM6, Kinmaze 등은 서로 다른 SNP를 가지고 있다고 하였다.
본 연구에서 RTSV 저항성 품종으로 선발된 품종 중에서 저항성반응이 상대적으로 강하면서, 조생종 품종인 상주벼에 대해 RTSV저항성 유전분석을 수행하였다. 감수성 품종인 운광벼와 저항성 품종인 상주벼가 교배된 F1 식물체는 모두 감수성이였다. 즉 상주벼의 RTSV 저항성 유전자는 열성유전자를 가지고 있는 것으로 판단되었다. 또한, 이들 F2 집단을 이용하여 RTSV 생물검정을 수행한 결과 저항성이 85개체, 이병성 116개체 분리하였으며, 이는 2개의 열성유전자의 이론적 분리비인 7 : 9에 적합하게 분리되었다. F1과 F2 집단의 유전분석 결과를 종합하면, 상주벼의 RTSV 저항성 유전자는 2개의 열성유전자에 의해 지배됨을 알 수 있었다(Table 3).
Table 3
Genetic analysis of rice tungro spherical virus (RTSV) in F2 population derived from a cross Unkwang and Sangju.
Table 3
Population RTSV resistance test Expected ratio X2 P-value
Resistant Susceptible Total
Unkwang/Sangju 85 116 201 7 : 9 0.174 0.676
F2
Imbe et al. (1995) RTSV저항성 유전분석 결과 Utri Merah82는 1개의 열성유전자 tsv-1을 가지고 있었고, Utri Merah80후대의 F3 집단 분석에서 2개의 독립적인 열성유전자 tsv-1과 tsv-2가 존재한다고 하였다. Azzam et al. (2001) Utri Merah80/TN1이 교배된 F1에서는 모두 감수성이었으며, 이들의 F3 집단에서 저항성, 분리 및 이병성이 각각 7 : 8 : 1로 분리되어 2개의 열성유전자를 가지고 있다고 것을 재확인하였다. 이는 본 연구의 상주벼에서 RTSV저항성 유전분리 양상과 유사한 결과를 나타내었다.
금후 상주벼의 저항성은 tsv1과 tsv2를 가진 품종과 대립성 검정 등을 통하여 동일한 유전자 유무에 대한 분석이 필요하다. 또한 상주벼의 RTSV 저항성 유전자에 대한 분자생물학적 연구를 지속적으로 수행하여, 퉁그로바이러스 저항성원의 다양화와 유전자집적(gene pyramiding)을 통한 저항성 안정화에 기여할 것으로 기대된다.
한국 자포니카 품종에 대한 퉁그로바이러스병 저항성 검정을 수행한 결과 RTSV에 이병율이 20% 이하로 저항성 반응을 보인 품종은 상주벼, 화성벼, 동진벼 이었으며, RTBV에는 모두 이병성이 있었다. RTSV 저항성 품종으로 선발된 품종 중에서 저항성반응이 상대적으로 강하면서, 조생종 품종인 상주벼에 대해 RTSV저항성 유전분석을 수행하였다. 감수성 품종인 운광벼와 저항성 품종인 상주벼가 교배된 F2 집단을 이용하여 RTSV 생물검정을 수행한 결과 저항성을 85개체, 이병성을 116개체 분리하였으며, 이는 2개의 열성유전자의 이론적 분리비인 7 : 9에 적합하게 분리되었다. 상주벼의 RTSV 저항성 유전자는 2개의 열성유전자에 의해 지배됨을 알 수 있었다.
본 성과물은(논문)은 농촌진흥청 연구사업(세부과제번호: PJ01319602)의 지원에 의해 이루어진 것임.
  • 1. Azzam O, Imbe T, Ikeda R, Nath PD, Coloquio E. 2001. Inheritance of resistance to rice tungro spherical virus in a near-isogenic line derived from Utri Merah and in rice cultivar TKM6. Euphytica 122: 91-97.
  • 2. Cabauatan PQ, Cabunagan RC, Koganezawa H. 1995. Biological variants of rice tungro viruses in the Philippines. Phytopathology 85: 77-81.
  • 3. Imbe T, Hashim H, Iwasaki M, Omura T. 1993. Resistance in some japonica rice cultivars to rice tungro spherical virus. Japan J Breed 43: 549-556.
  • 4. Imbe T. 1995. The development of stabilization technology for rice double cropping in the tropics. Final Report. IRRI Government of Japan Collaborative Research Project (December 1989 November 1994).
  • 5. Lee BC, Yoon YN, Hong SJ, Hong YK, Kwak DY, Lee JH, Yeo US, Kang HW, Hwang HG. 2008. Analysis on the occurrence of rice stripe virus. Res Plant Dis 14: 210-213.
  • 6. Lee JH, Muhsin M, Atienza GA, Kwak DY, Kim SM, DeLeon TB, Angeles D, Coloquio E, Kondoh H, Satoh K, Cabunagan RC, Cabauatan PQ, Kikuchi S, Leung H, Choi IR. 2010. Single nucleotide polymorphisms in a gene for translation initiation factor (eIF4G) of rice (Oryza sativa) associated with resistance to rice tungro spherical virus. Mol Plant Microbe Interact 23: 29-38.
  • 7. Hibino H, Saleh N, Roechan M. 1979. Transmission of two kinds of rice tungro associated viruses by insect vectors. Phytopathology 69: 1266-1268.
  • 8. Shim J, Torollo G, Angeles-Shim RB, Cabunagan RC, Choi IR, Yeo YS, Ha WG. 2015. Rice tungro spherical virus resistance into photoperiod-insensitive japonica rice by marker-assisted selection. Breed Sci 65: 345-351.

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Identification of Resistance to Rice Tungro Virus Disease in Korean Japonica Rice Cultivars
Korean. J. Breed. Sci.. 2019;51(2):86-90.   Published online June 1, 2019
DOI: https://doi.org/10.9787/KJBS.2019.51.2.86
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Identification of Resistance to Rice Tungro Virus Disease in Korean Japonica Rice Cultivars
Korean. J. Breed. Sci.. 2019;51(2):86-90.   Published online June 1, 2019
DOI: https://doi.org/10.9787/KJBS.2019.51.2.86
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Identification of Resistance to Rice Tungro Virus Disease in Korean Japonica Rice Cultivars
Identification of Resistance to Rice Tungro Virus Disease in Korean Japonica Rice Cultivars

Infection ratio of rice tungro spherical virus (RTSV) and rice tungro bacilliform virus (RTBV) in Korean Japonica varieties.

Entry No. Varieties RTBV RTSV Entry No. Varieties RTBV RTSV
1 Aranghyangchalbyeo 73.7 89.5 46 Jongnambyeo 100.0 37.8
2 Ansanbyeo 100.0 45.5 47 Juanbyeo 100.0 36.4
3 Anseongbyeo 100.0 83.3 48 Junganbyeo 100.0 27.5
4 Bongkwangbyeo 100.0 58.8 49 Junghwabyeo 100.0 62.5
5 Cheonmabyeo 100.0 27.8 50 Jungsangbyeo 100.0 65.8
6 Chucheongbyeo 100.0 73.0 51 Kwanganbyeo 100.0 63.2
7 Daeanbyeo 100.0 75.0 52 Mananbyeo 100.0 26.3
8 Daechangbyeo 100.0 38.5 53 Manchubyeo 100.0 62.5
9 Daejinbyeo 100.0 29.6 54 Mangeumbyeo 100.0 45.5
10 Daepyungbyeo 95.0 85.0 55 Manhobyeo 100.0 52.2
11 Daeribbyeo 1 100.0 41.2 56 Manmibyeo 100.0 25.9
12 Daesanbyeo 100.0 24.3 57 Manpungbyeo 100.0 41.0
13 Daeyabyeo 100.0 69.2 58 Mihyangbyeo 100.0 61.1
14 Dongjin1 100.0 55.0 59 Naepungbyeo 100.0 64.7
15 Dongjinbyeo 100.0 8.6 60 Nampyeongbyeo 100.0 54.3
16 Dongjinchalbyeo 100.0 51.4 61 Namweonbyeo 87.1 45.2
17 Dunnaebyeo 100.0 26.7 62 Nonganbyeo 100.0 34.3
18 Gancheokbyeo 100.0 42.5 63 Nongbaek 100.0 47.2
19 Geumanbyeo 100.0 54.5 64 Obongbyeo 100.0 26.7
20 Geumnambyeo 100.0 34.3 65 Odaebyeo 100.0 61.1
21 Geumobyeo 87.2 46.2 66 Palgongbyeo 100.0 52.9
22 Geumobyeo 100.0 76.2 67 Saegyehwabyeo 84.6 61.5
23 Goamy 100.0 62.5 68 Sambaekbyeo 100.0 37.0
24 Gurubyeo 100.0 78.8 69 Samcheonbyeo 100.0 28.2
25 Haepyeongbyeo 100.0 33.3 70 Sangjubyeo 94.1 0.0
26 Heughyangbyeo 100.0 47.2 71 Sangmibyeo 100.0 64.3
27 Heugjinjubyeo 97.1 37.1 72 Sangnambadtbyeo 89.2 48.6
28 Heugnambyeo 100.0 55.0 73 Sanjuchalbyeo 100.0 44.1
29 Hoanbyeo 100.0 25.0 74 Seoanbyeo 100.0 28.6
30 Hwaanbyeo 100.0 44.4 75 Seojinbyeo 87.9 60.6
31 Hwabongbyeo 100.0 63.2 76 Seolgengbyeo 100.0 55.3
32 Hwadongybeo 100.0 21.1 77 Seolhyangchalbyeo 100.0 32.4
33 Hwajinbyeo 100.0 63.6 78 Seomjinbyeo 100.0 35.0
34 Hwajungbyeo 100.0 72.2 79 Sinseonchalbyeo 100.0 30.8
35 Hwamyeongbyeo 100.0 62.9 80 Sobaekbyeo 100.0 53.6
36 Hwanambyeo 100.0 91.7 81 Sobibyeo 100.0 80.0
37 Hwasambyeo 100.0 61.5 82 Sujinbyeo 100.0 41.7
38 Hwaseongbyeo 92.1 7.9 83 Taeseongbyeo 94.3 62.9
39 Ilmibyeo 92.5 25.0 84 Tamjinbyeo 100.0 65.2
40 Ilpumbyeo 100.0 48.6 85 Umbongbyeo 100.0 45.7
41 Janganbyeo 100.0 21.6 86 Undoobyeo 73.7 78.9
42 Jinbongbyeo 100.0 58.3 87 Unjangbyeo 100.0 35.3
43 Jinbuolbyeo 100.0 37.8 88 Weonhwangbyeo 100.0 38.5
44 Jinheung 100.0 82.4 89 Yeonganbyeo 100.0 35.7
45 Jinjubyeo 100.0 68.0

The infection ratio of moderate resistant varieties against to rice tungro spherical virus (RTSV).

Varieties Infection ratio (%)
RTBV+RTSV RTBVz RTSV
Sangjubyeo 0.0 94.1 0.0
Hwaseongbyeo 7.9 84.2 0.0
Dongjinbyeo 8.6 85.7 0.0

zRice tungro bacilliform virus

Genetic analysis of rice tungro spherical virus (RTSV) in F2 population derived from a cross Unkwang and Sangju.

Population RTSV resistance test Expected ratio X2 P-value
Resistant Susceptible Total
Unkwang/Sangju 85 116 201 7 : 9 0.174 0.676
F2
Table 1 Infection ratio of rice tungro spherical virus (RTSV) and rice tungro bacilliform virus (RTBV) in Korean Japonica varieties.
Table 2 The infection ratio of moderate resistant varieties against to rice tungro spherical virus (RTSV).

Rice tungro bacilliform virus

Table 3 Genetic analysis of rice tungro spherical virus (RTSV) in F2 population derived from a cross Unkwang and Sangju.

Table 1

Table 2

Table 3

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