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잎이 얇고 작은 구를 형성하는 배추 계통 ‘원교20042호’ 육성

Developing Inbred Lines of ‘Wonkyo20042ho’ Bearing Thin Leaves and Small Sized Head in Kimchi Cabbage (Chinese Cabbage) (Brassica rapa. L)

Ha-Young Jang1, Myeong-Whoon Seo2, Min Young Park1, Moo-Kyoung Yoon3, Jung-Soo Lee4, Suhyoung Park1,*
The Korean Journal of Breeding Science 2016;48(2):184-191.
Published online: May 31, 2016

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*Corresponding author (psh@korea.kr, +82-63-238-6622, +82-63-238-6605)
• Received: October 7, 2015   • Accepted: January 22, 2016

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  • Kimchi cabbage (known as Chinese cabbage) is one of the major vegetable in Korea. An inbred line ‘Wonkyo20042ho’, was developed by the National Institute of Horticultural & Herbal Science, RDA in South Korea. The first cross carried out using ‘Norang gwandong’ which had been introduced to Korea in 2007. During segrerating generations, bud pollination carried out using selected plants to fix genetic materials. ‘Wonkyo20042ho’ inbred line was finally selected by breeders from commercial companies at the field exhibition at 2011. The horticultural traits of ‘Wonkyo20042ho’ inbred line were as follows; half-open head, yellow inner leaf colour, and dark green outer leaf colour. The average head weight was 1899.2g, head width and height were 17.5cm and 27.9cm, respectively. Numbers of outer and internal leaves were 14.3 and 66.5, respectively. Harvest maturity was mid-late maturing type and average of seed yield for three years was 723 seeds. In addition, ‘Wonkyo20042ho’ inbred line showed resistance to clubroot disease (race 4 single spore isolation) and possessed thin leaves and small sized head.
배추(Brassica rapa L.)는 한국의 주요 채소로 고려시대인 13세기경부터 이용되어 왔으며 김치의 주 재료로 한국인의 사랑 을 받고 있다. 2014년도에는 1인당 연간 소비량이 58.5kg로 꾸준히 증가하는 추세를 보이며, 최근 김치의 우수성이 국제적으 로 입증되며 식습관이 유사한 중국, 일본은 물론 미주와 유럽 국가에 까지 소비가 증가하고 있다.
국내 배추 재배면적은 연간 차이는 있지만 2014년도에 약 32만 ha이며, 연간 생산량은 약 25만 톤으로 우리나라 주요 5대 채소에 포함된다(MAFRA 2015). 또한 한국 배추 품종의 우수성이 인정되어 중국, 일본, 동남아 등에 2.8백만 US$를 수출하여 채소 종자 수출액의 7%를 차지하였다(Korea seed association 2014).
종자 수출 패턴을 살펴보면 5년전인 2010년에는 2백만 US$로 전체 수출액의 9%, 2006년 1.5백만 US$로 전체 수출액의 8%를 차지하였다. 지난 10년 동안 종자 수출 금액은 소폭 증가하였으 나 전체 채소종자 수출액에서 차지하는 비중은 오히려 소폭 감소하였으며 이는 주요 수출 국가인 중국의 배추 품종 개발 노력 등에 의해 가격 경쟁이 발생하였기 때문으로 해석된다. 따라서 한국 배추 품종의 부가가치를 향상시킬 수 있도록 병에 저항성이며 원예적 특성이 양호한 계통의 육성이 필요하게 되었 다.
뿌리혹병은 절대 활물 기생균인 Plasmodiophora brassicae에 의해 발생하는 토양전염성 병으로(Braselton 1995), 처음엔 진 균(fungi)으로 분류되었으나 DNA 염기서열 분석 결과 진균과는 연관이 없는 것으로 밝혀졌으며(Castlebury & Domier 1998, Down et al. 2002), ribosomal RNA의 발생학적 분류 결과 원생동물(protozoa)로 분류되었다(Van de Peer et al. 2000). 현재 뿌리혹병은 우리나라 배추의 모든 재배 작형에 전국적으로 발병하고 있으며, 발병 시 수량의 30~70% 정도를 감소시킴으로 배추 재배에 큰 위협을 주고 있다. 따라서 뿌리혹병의 방제 방법 및 저항성 품종 개발 관련 연구가 지속적으로 추진되고 있다. 뿌리혹병 방제를 위해 화학 약품, 농약 및 길항균을 이용하는 방법이 개발되었으나(Datnoff et al. 1987, Cheah et al. 2000, Yeoung et al. 2003), 처리 비용(약제비 및 노동비)이 많이 소모 되는 문제가 있다. 또한 방제 효과도 처리한 연도의 재배에는 효과가 있으나 이듬해 재배할 경우 휴면포자 상태로 생존한 병원균의 밀도가 높아지면서 병이 다시 발생하는 문제가 존재한 다. 나아가 최근에는 저항성 품종 개발을 위하여 뿌리혹병 저항성 자원인 순무를 이용해 배추와 교배하여 소포자 배양을 활용해 비교적 빠른 시간 내에 뿌리혹병 저항성 배추 계통을 육성하기도 하였다(Park et al. 2011).
국립원예특작과학원 채소과에서는 뿌리혹병에 저항성을 갖 는 다양한 배추 계통 육성을 위해 국내외 여러 품종을 수집하여 육종 재료로 이용하여 병에도 저항성이며 원예적 특성도 우수한 계통을 육성코자 노력하였다. 또한 육성된 계통의 민간 활용도를 높이기 위해 포장 공개 전시회 과정을 거쳐 다양한 민간 육종가의 의견을 수렴하여 공동으로 선발하였다.
‘원교20042호’는 2007년까지 채소과에서 수집한 한국 배추 시판품종 중하나인 ‘노랑관동’에 뿌리혹병 race4를 인공접종하 여 살아남은 개체를 뇌수분하여 육성하였다. ‘노랑관동’은 홍농 종묘에서 육성한 시판품종으로 고랭지 여름재배용 품종으로 겉잎은 진한 녹색, 속잎은 노랑색으로 맛이 우수하며 증룩이 얇은 엽수형 품종이다.
뿌리혹병 인공 접종법
각 품종 별로 종자 30립을 9cm petridish에 여과지 2장을 깔고 파종한 후 물이 마르지 않도록 관리하며 발아를 시켰다. 파종 온도에 따라 떡잎 전개 시기가 다르지만 대체로 7~9일이 되어 떡잎이 완전하게 펼쳐지면 단포자 분리로 획득된 race4 (Kang et al. 2000)를 증류수에 갈은 후 약 5 x 105 농도로 계산된 접종액으로 접종하였다. 접종 1일 이후 상토를 채운 50공 프러그 트레이에 정식한 후 접종액이나 관주 양액이 밖으로 흘러나가 주변을 오염시키지 않도록 트레이 물받침을 받혀주었 다. 재배 시 관수 용액이 밖으로 흐르지 않도록 프라스틱 상자로 화분을 받쳐주었다.
뿌리혹병 저항성 평가법
육묘용 한방 양액으로 9주간 재배 후 뿌리를 조사하였다. 뿌리혹병 발생 정도에 따라 병증을 보이지 않는 개체수부터 병증이 가장 심한 개체수까지 무증상의 1부터 중증의 5로 단계를 나눈 후(Fig. S1) 조사하여 아래의 공식에 대입하여 DI 값을 산출했다(Werner et al. 2008).
Fig. S1.
Kimchi cabbage roots after infection with P. brassicae. The roots are arranged from left to right according to the degree of distorted root growth. The clubroot disease rating scale ranges between 1 (left, resistant) and 5 (right, fully sensitive).
KJBS-48-2-184_FS1.gif
Disease Index(DI)={(1n1 + 2n2 + 3n3 + 4n4 + 5n5) × 100}/ (4 x NT)
또한, 뿌리에 혹을 형성하지 않은 개체는 소포자 배양을 위해 지름 10cm의 소형 화분에 옮겨 심었다.
뇌수분에 의한 종자 증식
뇌수분을 위한 식물체는 춘화처리(5°C, 10주간)한 후 온실로 옮겨 지름 20cm의 화분에 정식하여 개화를 유도하였다. 이식 후 추대가 발생하면 1차 추대는 제거하여 여러 개의 2차 추대를 유도한 후 지지대를 세워서 추대가 꺾이지 않도록 관리하면서 교배를 실시하였다. 뇌수분은 꽃봉오리의 길이가 5mm정도로 아직 개화하지 않은 봉오리의 윗부분을 제거하고 봉지 안에서 개화된 동일한 개체의 화분을 이용하여 수분한 후 다시 일주일간 봉지를 씌워 오염 수분을 방지하였다. 뇌수분 후 50~60일이 지나 종자가 노랗게 익으면 꼬투리가 벌어지기 전에 채종하였다.
소규모 망실을 이용한 채종 실험
채종 실험을 위한 식물체는 50공 트레이에 파종한 후 본엽이 3매정도 자랐을 때 저온 챔버로 옮겨 춘화처리(5°C, 10주간)하였 다. 이후 가로 세로 1m x 1m 크기의 망실에 4~6주씩 정식하였다. 정식 후 추대가 발생하면 1차 추대는 제거하여 여러 개의 2차 추대를 유도한 후 지지대를 세워서 추대가 꺾이지 않도록 관리하 면서 채종 실험을 실시하였다. 뒤영벌의 수분 활동을 활발하게 하기 위해, 모든 개체가 개화한 이후 한꺼번에 꽃대를 전정하여 망실 내부 개체의 개화 시기를 동일하게 조절한 후 벌을 투입하였 다. 1차 투입 후 15일이 경과한 시점에 2차로 벌을 재투입 하였으 며, 2차 투입 후 20일 경과 후 망을 제거하고 화지를 정리하여 재배관리 하였다. 이후 꼬투리가 청색에서 황색으로 바뀌는 시점 에 종자를 수확하여 채종 및 정선을 실시하였다.
원예적 특성 평가
종자가 성공적으로 증식된 육성된 계통의 원예적 특성을 평가 하기 위해 수원시 장안구 이목동 소재 국립원예특작과학원 채소 과 노지 포장에서 가을에 재배하였다. 재배 방법은 배추 표준재배 법에 준하여 재배하였으며, 검은색 멀칭 필름으로 피복 하고 점적관수를 실시하였다. 파종은 8월 1일~5일, 정식은 8월 28 일~9월 4일에 행했다. 특성조사는 국립종자원의 배추 특성조사 요령(UPOV 2003)과 농촌진흥청 시험 연구 설계서의 조사기준 에 준하여 11월 1일~20일 사이에 실시하였다.
육성경위
배추 ‘원교20042호’는 국내 시판품종인 ‘노랑관동’의 뿌리혹 병 저항성 선발개체를 뇌수분으로 세대진전하여 육성된 계통으 로(Fig. 1) 결구 형태는 반포합형이며, 속잎이 노랗고 잎이 얇아 맛이 좋은 중간모본이다. 2007년까지 채소과에서 수집한 국내 시판 종 50품종(Table 1)을 뿌리혹병 race4균주로 접종하여 생존한 26품종의 저항성 선발 개체 133점을 소형 화분에 정식하 였다(Table 2). 정식된 개체들은 뇌수분을 통하여 종자를 증식하 였으나, 대부분의 개체는 종자 증식 중간에 병증이 발생하는 등 종자의 증식이 불가능하였다. 이후 성공적으로 증식된 개체의 종자를 활용하여 뿌리혹병 저항성을 조사하여 선발을 실시하였 다.
Fig. 1.
Pedigree diagram of ‘Wonkyo20042ho’
KJBS-48-2-184_F1.gif
Table 1.
Collection list of domestic commercial cultivars of Kimchi cabbagez) up to 2007.
Table 1.
No. Introduction year Cultivar Company
1 1993 Gonaengji yeoreum baechu Jungang
2 1993 Gonaengji josaengyeoreum baechu Konong
3 1993 Jangsu yeoreum Nongwoo
4 1993 Tongilsagye baechu Hannong
5 1993 93-2-1 Kimphilju
6 2001 Danmat baechu Heungnong
7 2001 Jangmi baechu Nongwoo
8 2001 Gyeoul norang baechu Sakada
9 2001 Geumbangul baechu Nongwoo
10 2001 Naeseogarak baechu Jungang
11 2001 Yeongnong1 baechu Kyoungshin
12 2001 Buram3 Heungnong
13 2001 Noranja baechu Nongwoo
14 2001 House geumgarak(2) Jungang
15 2001 Chilseong baechu Nongwoo
16 2001 Gangnyeok yeoreum(2) Heungnong
17 2001 Saennorang baechu(11) Nongwoo
18 2004 Ssammat baechu(2) Nongwoo
19 2004 Chilseong baechu (2) Nongwoo
20 2004 Noranja baechu Nongwoo
21 2004 Tambok baechu Nongwoo
22 2004 Asia mini baechu (3) Asia
23 2004 Asia Norangmini(2) Asia
24 2004 Jinnorang baech Seoul
25 2004 Medakkassam baechu Takii
26 2004 Pyeongseongtong baechu PECR
27 2004 Pyeongseongtong baechu PECR
28 2005 Baechu55 Nongwoo
29 2005 CR Geurin Nongwoo
30 2005 Jinhangeum Heungnong
31 2005 Norang gimchi Hannong
32 2005 Jeongilpum Heungnong
33 2005 Sinhawang baechu Jungang
34 2005 Josaenggarak baechu Heungnong
35 2005 Bokdori baechu Heungnong
36 2006 Tamna baechu(2) Heungnong
37 2006 Yeppeun noraengi baechu(5) Nongwoo
38 2006 CR Matgeurin baechu Jungang
39 2006 Samjin baechu(2) Heungnong
40 2006 Dongpum baechu(2) Nongwoo
41 2006 Ganghan baechu(2) Sakada
42 2006 CR Myeongpum baechu Heungnong
43 2006 Geumgarak baechu Jungang
44 2006 Norang ssam baechu Sakada
45 2006 Okhwang ssial baechu Sakada
46 2007 Chaempion baechu(3) Heungnong
47 2007 Sseongeurin(3) Heungnong
48 2007 Buram peulleoseu baechu(3) Heungnong
49 2007 Jincheong baechu(3) Heungnong
50 2007 Norang gwandong baechu(3) Heungnong

z)Kimchi cabbage is the name of a subgroup of Chinese cabbage which is used as a ingredient of Korean traditional cuisine, Kimchi (codex, 2012).

Table 2.
Result of screening for clubroot inoculation using commercial cultivar of Kimchi cabbage.
Table 2.
No. Resistance (No.) Medium sensitive (No.) Sensitive (No.) Total (No.) Resistance percentage (%)
7 14 14 0 28 50
8 0 2 0 2 0
9 11 7 0 18 61.1
10 14 9 0 23 60.9
11 0 10 0 10 0
12 9 6 0 15 60
13 0 6 0 6 0
14 1 3 0 4 25
15 5 10 0 15 33.3
16 7 3 0 10 70
17 1 2 0 3 33.3
18 2 14 0 16 12.5
19 2 3 0 5 40
20 2 10 0 12 16.7
21 1 0 0 1 100
22 0 5 0 5 0
23 1 4 0 5 20
25 0 9 0 9 0
27 7 4 0 11 63.6
28 6 12 0 18 33.3
30 3 19 0 22 13.7
32 0 2 0 2 0
33 1 3 0 4 25
36 1 11 0 12 8.3
37 0 10 0 10 0
38 4 9 0 13 30.8
39 1 2 0 3 33.3
40 12 10 0 22 54.5
41 2 2 0 4 50
42 0 5 0 5 0
43 0 3 0 3 0
44 0 1 0 1 0
45 1 13 0 14 7.1
46 0 20 0 20 0
47 3 16 0 19 15.8
48 0 1 0 1 0
49 9 16 0 25 36
50 13 10 0 23 56.5

z)The number was represented equal cultivar in table 1. Some cultivar was not germinated.

y)CR test was performed using a single spore isolate with race 4.

x)Resistance; lv. 1, medium sensitive; lv. 2~3 , sensitive; lv. 4~5 (in Fig. S1)

최종 선발 개체인 ‘노랑관동’의 후대는 뇌수분으로 종자를 증식한 후 생육 특성을 조사하였다. 원예적 특성이 고정되어 순도가 높은 것으로 확인된 계통들은 2011년 가을에 종묘회사와 공동으로 선발하였다. 최종적으로 선발된 계통은 그 종자를 증식 함과 동시에 직무육성신품종선정심의회에 상정하였고, 민관 채 소 육종가 및 연구자의 동의로 우수한 성적으로 통과되었다. 이후 2차례의 종자 증식 과정을 거친 후 2014년에 ‘원교20042 호’로 명명하여 품종보호출원(출원2014-582)하였다.
생육 및 수량특성
‘원교20042호’의 초자는 중간이며, 결구형은 반포합형으로 속잎색이 노랗고 코어 부분이 작은 결구를 형성한다. 겉잎색은 초록색이며 잎의 모양은 계란형으로 모용은 중간 정도이며 숙기 도 중간으로 대비종과 유사하다(Table 3, Fig. 2). ‘원교20042호’ 의 수량 및 가변특성은 시비량에 의해 변동이 있었으나, 대체적으 로 대조인 ‘불암3호’보다 작았다(Table 4).
Table 3.
Morphological characteristics of ‘Wonkyo20042ho’.
Table 3.
Cultivar Plant: height Head
 Leaf
 Harvest maturity Outer leaf Head internal
Type Outer color z) Internal color y) Shape No. of blisters on upper side
Wonkyo20042ho Medium Half-open 3.9 4.2 Ovate Medium Medium
Buram3 Medium Half-open 4.2 4.1 Ovate Medium Medium

z)Color chart: 0; DIC 2087, 1; DIC 2088, 2; DIC 2089, 3; DIC 2540, 4; DIC 167, 5; DIC 166, 6; DIC 86, 7; DIC 85, 8; DIC 84

y)Color chart: 0; DIC 361, 1; DIC 362, 2; DIC 246, 3; DIC 369, 4; DIC 2324, 5; DIC 2336, 6; DIC 2325, 7; DIC 2356, 8; DIC 2355

Fig. 2.
Inner shape of ‘‘Wonkyo20042ho’ inbred line planting bud pollinated seeds at 2011, and 1st and 2nd reproduced seeds using bee pollination at 2012, 2013, respectively.
KJBS-48-2-184_F2.gif
‘원교20042호’ 식물체 전체 무게는 3,153.2g으로 대조의 4,349.7g보다 가벼웠으며, 구의 무게도 1,899.2g으로 대조의 3,003.0g보다 가벼웠다. 구의 폭은 17.5cm로 대조의 21.2cm보 다 좁았으나, 구의 높이는 27.9cm로 대조의 27.5cm보다 길어 다소 좁고 긴 형태의 구를 형성하였다. ‘원교20042호’의 잎의 길이는 47.4cm로 대조인 ‘불암3호’의 49.7cm보다 짧았으며, 잎의 폭도 29.7cm로 대조의 34.8cm보다 좁았다. 겉잎의 개수는 14.3개로 대조의 13.7개보다 많았으며, 구를 형성하고 있는 속잎 의 개수는 66.0개로 대조의 75.1개보다 적었다(Table 4). ‘원교 20042호’의 수량성은 43.9t/ha으로 대조보다 적었으나, 구와 잎의 특성은 중간모본으로서는 우수한 것으로 포장품평회(’10) 에서 농우바이오 등 4개사에서 선발되었다.
Table 4.
Variable characteristics of ‘Wonkyo20042ho’.
Table 4.
Cultivar Plant weight (g) Head
 Leaf
Weight (g) Width (cm) Height (cm) Outer length (cm) Outer width (cm) No. outer No. inner
Wonkyo20042ho 3153.2±440.4 1899.2±239.3 17.5±0.9 27.9±1.06 47.4±2.5 29.7±1.4 14.3±2.7 66±5.6
Buram3 4349.7±1002.9 3003±869.6 21.2±1.6 27.5±2.0 49.7±2.4 34.8±3.2 13.7±0.4 75.1±1.4

z)Values are the mean ±SE (n=3, 3 years are 2012, 2013, 2014).

뿌리혹병 저항성
우리나라 모든 배추 재배 지역과 작형에 고르게 발생하여 수량 감소를 야기하는 주요 병인 뿌리혹병 저항성의 인공 접종 결과, ‘원교20042호’는 뿌리혹병 race4에 저항성을 보여 DI(Disease Index)가 3.1로 대조인 춘광의 21보다 훨씬 낮았다 (Table 5). 이는 대부분 육성 계통의 DI 값이 125이고, 중국도입 종들의 DI 값이 100~125 사이에 분포하고 있는 것에 비하면 원교20042호는 뿌리혹병 race4에 매우 저항성임을 알 수 있었다.
Table 5.
Disease index(DI) z) of ‘Wonkyo20042ho’.
Table 5.
Cultivar DI
Wonkyo20042ho 3.1
Chungwang 21.0
04-1-21 (Cultivar introduced from China) 108.3
04-1-22 (Cultivar introduced from China) 121.4
04-1-23 (Cultivar introduced from China) 121.4
06-1-24 (Cultivar introduced from China) 125.0
06-1-25 (Cultivar introduced from China) 100.0
09-FK74-3 (Breeding line) 62.5
09-FK267-2 (Breeding line) 125.0
09-FK268-1 (Breeding line) 125.0
09-FK281-2 (Breeding line) 125.0
09-FK283-1 (Breeding line) 125.0
09-FK284-1 (Breeding line) 125.0

zDisease Index (DI) = {(1n1 + 2n2 + 3n3 + 4n4 + 5n5) x 100}/ (4 x NT) n1; Resistance plant, n2~3; Medium sensitive plant, n4~5; Sensitive plant, NT; Total plant

채종특성
‘원교20042호’의 주당 채종량은 년도별로 차이가 있어 2012 년도에는 1586.1립, 2014년도에는 227.6립, 2015년도에는 335.4립이었다(Table 6). 이렇게 년도별로 채종량에 큰 차이를 보이는 이유는 강우빈도에 따른 것으로 추정해 볼 수 있다. 기상 청 자료를 분석해보니(수원 ’12, ’14, 전주’15) 종자가 익어가는 시기인 6월에 2014년도와 2015년도에는 강우빈도가 빈번했으 나, 2012년에는 초순부터 중순까지 전혀 비가 오지 않아 종자가 익는데 좋은 환경이 형성된 것으로 사료된다. 최근 콩과작물인 헤어리베치에서 비슷한 연구가 수행되었는데 개화 후 42~54일 기간 내에 장마를 회피할 수 있는 수확기간이 헤어리베치 다수확 채종 기간으로 판단된다는 결론을 내렸다(Ku et al. 2012). 이는 기존의 육종상식과 일치하는 것으로 종자가 익는 시기에 강우빈 도가 중요하다는 것을 나타낸다. 이와 더불어 배추에서 다양한 환경스트레스에 관련한 연구도 활발히 진행되고 있으며 최근에 BrTSR35라는 유전자가 배추의 저온, 고농도의 염과 건조 등의 환경스트레스에 대한 저항성 유전자로 밝혀졌다(Kim et al. 2014). 이러한 정보를 활용하여 농작물의 환경스트레스 저항성 을 높여줄 수 있는 주요한 유전자원으로 이용하면 작물의 생산성 을 높일 수 있을 것으로 사료된다.
Table 6.
Seed production of ‘Wonkyo20042ho’ using small net house in 3 years (’12, ’14, and ’15).
Table 6.
Cultivar Location Year Seeds per plant (No.)
Wonkyo20042ho Suwon 2012 1586.1±723
2014 227.6±62.5
Wanju 2015 355.4±46.1

zValues are the mean ±SE (n=3) for each year.

재배상 유의점
‘원교20042호’는 중간모본으로 육성된 계통으로 일대잡종 품종과 달리 환경에 민감하며 특히 고온·건조 조건에서 석회 결핍이 잘 발생되므로 퇴비와 비료가 충분한 포장에서 관수에 유의하며 재배해야 한다.
배추(Kimchi cabbsage)는 한국의 주요 채소 중 하나이다. ‘원교 20042호’는 농촌진흥청 국립원예특작과학원에서 2007년에 국내 시판품종인 ‘노랑관동’의 뿌리혹병 저항성 선발개체를 뇌수분으 로 세대진전하여 육성된 계통이다. 최종선발을 위해 종묘회사와 공동으로 포장 품평회를 실시하여(’11) ‘0750-8-1-1’을 선발하였 다. 선발 계통은 직무육성신품종 심의를 거쳐 2014년에 ‘원교 20042호’로 품종보호출원 하였다. ‘원교20042호’의 원예적 특 성으로는 결구형은 반포합형으로 속잎색이 노랗고 겉잎색은 초록색이다. 또한 구중은 1,899.2g으로 구의 폭과 높이는 각각 17.5cm, 27.9cm로 다소 좁은 구를 형성한다. 겉잎과 속잎의 수는 각각 14.3개, 66.5개 이며, 숙기는 중간으로 3년 평균 채종량 이 723개 이다. 더욱이 ‘원교20042호’는 뿌리혹병 race4에 저항 성을 보이며 잎이 얇고 구가 작은 특성을 갖는다.
본 논문은 농촌진흥청 연구사업(과제번호: PJ010275012016)와 농림수산식품부 GSP 채소종자 사업단의 연구과제(과제번 호: 213002-04-3-SB120)의 지원에 의해 이루어졌습니다. 또한 본 연구는 2016년도 농촌진흥청 국립원예특작과학원 박사후연 수과정지원사업에 의해 이루어졌습니다.
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Developing Inbred Lines of ‘Wonkyo20042ho’ Bearing Thin Leaves and Small Sized Head in Kimchi Cabbage (Chinese Cabbage) (Brassica rapa. L)
Korean. J. Breed. Sci.. 2016;48(2):184-191.   Published online June 30, 2016
DOI: https://doi.org/10.9787/KJBS.2016.48.2.184
Download Citation

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Developing Inbred Lines of ‘Wonkyo20042ho’ Bearing Thin Leaves and Small Sized Head in Kimchi Cabbage (Chinese Cabbage) (Brassica rapa. L)
Korean. J. Breed. Sci.. 2016;48(2):184-191.   Published online June 30, 2016
DOI: https://doi.org/10.9787/KJBS.2016.48.2.184
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Developing Inbred Lines of ‘Wonkyo20042ho’ Bearing Thin Leaves and Small Sized Head in Kimchi Cabbage (Chinese Cabbage) (Brassica rapa. L)
Image Image Image
Fig. S1. Kimchi cabbage roots after infection with P. brassicae. The roots are arranged from left to right according to the degree of distorted root growth. The clubroot disease rating scale ranges between 1 (left, resistant) and 5 (right, fully sensitive).
Fig. 1. Pedigree diagram of ‘Wonkyo20042ho’
Fig. 2. Inner shape of ‘‘Wonkyo20042ho’ inbred line planting bud pollinated seeds at 2011, and 1st and 2nd reproduced seeds using bee pollination at 2012, 2013, respectively.

Fig. S1.

Fig. 1.

Fig. 2.

Developing Inbred Lines of ‘Wonkyo20042ho’ Bearing Thin Leaves and Small Sized Head in Kimchi Cabbage (Chinese Cabbage) (Brassica rapa. L)

Collection list of domestic commercial cultivars of Kimchi cabbagez) up to 2007.

No. Introduction year Cultivar Company
1 1993 Gonaengji yeoreum baechu Jungang
2 1993 Gonaengji josaengyeoreum baechu Konong
3 1993 Jangsu yeoreum Nongwoo
4 1993 Tongilsagye baechu Hannong
5 1993 93-2-1 Kimphilju
6 2001 Danmat baechu Heungnong
7 2001 Jangmi baechu Nongwoo
8 2001 Gyeoul norang baechu Sakada
9 2001 Geumbangul baechu Nongwoo
10 2001 Naeseogarak baechu Jungang
11 2001 Yeongnong1 baechu Kyoungshin
12 2001 Buram3 Heungnong
13 2001 Noranja baechu Nongwoo
14 2001 House geumgarak(2) Jungang
15 2001 Chilseong baechu Nongwoo
16 2001 Gangnyeok yeoreum(2) Heungnong
17 2001 Saennorang baechu(11) Nongwoo
18 2004 Ssammat baechu(2) Nongwoo
19 2004 Chilseong baechu (2) Nongwoo
20 2004 Noranja baechu Nongwoo
21 2004 Tambok baechu Nongwoo
22 2004 Asia mini baechu (3) Asia
23 2004 Asia Norangmini(2) Asia
24 2004 Jinnorang baech Seoul
25 2004 Medakkassam baechu Takii
26 2004 Pyeongseongtong baechu PECR
27 2004 Pyeongseongtong baechu PECR
28 2005 Baechu55 Nongwoo
29 2005 CR Geurin Nongwoo
30 2005 Jinhangeum Heungnong
31 2005 Norang gimchi Hannong
32 2005 Jeongilpum Heungnong
33 2005 Sinhawang baechu Jungang
34 2005 Josaenggarak baechu Heungnong
35 2005 Bokdori baechu Heungnong
36 2006 Tamna baechu(2) Heungnong
37 2006 Yeppeun noraengi baechu(5) Nongwoo
38 2006 CR Matgeurin baechu Jungang
39 2006 Samjin baechu(2) Heungnong
40 2006 Dongpum baechu(2) Nongwoo
41 2006 Ganghan baechu(2) Sakada
42 2006 CR Myeongpum baechu Heungnong
43 2006 Geumgarak baechu Jungang
44 2006 Norang ssam baechu Sakada
45 2006 Okhwang ssial baechu Sakada
46 2007 Chaempion baechu(3) Heungnong
47 2007 Sseongeurin(3) Heungnong
48 2007 Buram peulleoseu baechu(3) Heungnong
49 2007 Jincheong baechu(3) Heungnong
50 2007 Norang gwandong baechu(3) Heungnong

Kimchi cabbage is the name of a subgroup of Chinese cabbage which is used as a ingredient of Korean traditional cuisine, Kimchi (codex, 2012).

Result of screening for clubroot inoculation using commercial cultivar of Kimchi cabbage.

No. Resistance (No.) Medium sensitive (No.) Sensitive (No.) Total (No.) Resistance percentage (%)
7 14 14 0 28 50
8 0 2 0 2 0
9 11 7 0 18 61.1
10 14 9 0 23 60.9
11 0 10 0 10 0
12 9 6 0 15 60
13 0 6 0 6 0
14 1 3 0 4 25
15 5 10 0 15 33.3
16 7 3 0 10 70
17 1 2 0 3 33.3
18 2 14 0 16 12.5
19 2 3 0 5 40
20 2 10 0 12 16.7
21 1 0 0 1 100
22 0 5 0 5 0
23 1 4 0 5 20
25 0 9 0 9 0
27 7 4 0 11 63.6
28 6 12 0 18 33.3
30 3 19 0 22 13.7
32 0 2 0 2 0
33 1 3 0 4 25
36 1 11 0 12 8.3
37 0 10 0 10 0
38 4 9 0 13 30.8
39 1 2 0 3 33.3
40 12 10 0 22 54.5
41 2 2 0 4 50
42 0 5 0 5 0
43 0 3 0 3 0
44 0 1 0 1 0
45 1 13 0 14 7.1
46 0 20 0 20 0
47 3 16 0 19 15.8
48 0 1 0 1 0
49 9 16 0 25 36
50 13 10 0 23 56.5

The number was represented equal cultivar in table 1. Some cultivar was not germinated.

CR test was performed using a single spore isolate with race 4.

Resistance; lv. 1, medium sensitive; lv. 2~3 , sensitive; lv. 4~5 (in Fig. S1)

Morphological characteristics of ‘Wonkyo20042ho’.

Cultivar Plant: height Head
 Leaf
 Harvest maturity Outer leaf Head internal
Type Outer color z) Internal color y) Shape No. of blisters on upper side
Wonkyo20042ho Medium Half-open 3.9 4.2 Ovate Medium Medium
Buram3 Medium Half-open 4.2 4.1 Ovate Medium Medium

Color chart: 0; DIC 2087, 1; DIC 2088, 2; DIC 2089, 3; DIC 2540, 4; DIC 167, 5; DIC 166, 6; DIC 86, 7; DIC 85, 8; DIC 84

Color chart: 0; DIC 361, 1; DIC 362, 2; DIC 246, 3; DIC 369, 4; DIC 2324, 5; DIC 2336, 6; DIC 2325, 7; DIC 2356, 8; DIC 2355

Variable characteristics of ‘Wonkyo20042ho’.

Cultivar Plant weight (g) Head
 Leaf
Weight (g) Width (cm) Height (cm) Outer length (cm) Outer width (cm) No. outer No. inner
Wonkyo20042ho 3153.2±440.4 1899.2±239.3 17.5±0.9 27.9±1.06 47.4±2.5 29.7±1.4 14.3±2.7 66±5.6
Buram3 4349.7±1002.9 3003±869.6 21.2±1.6 27.5±2.0 49.7±2.4 34.8±3.2 13.7±0.4 75.1±1.4

Values are the mean ±SE (n=3, 3 years are 2012, 2013, 2014).

Disease index(DI) z) of ‘Wonkyo20042ho’.

Cultivar DI
Wonkyo20042ho 3.1
Chungwang 21.0
04-1-21 (Cultivar introduced from China) 108.3
04-1-22 (Cultivar introduced from China) 121.4
04-1-23 (Cultivar introduced from China) 121.4
06-1-24 (Cultivar introduced from China) 125.0
06-1-25 (Cultivar introduced from China) 100.0
09-FK74-3 (Breeding line) 62.5
09-FK267-2 (Breeding line) 125.0
09-FK268-1 (Breeding line) 125.0
09-FK281-2 (Breeding line) 125.0
09-FK283-1 (Breeding line) 125.0
09-FK284-1 (Breeding line) 125.0

Disease Index (DI) = {(1n1 + 2n2 + 3n3 + 4n4 + 5n5) x 100}/ (4 x NT) n1; Resistance plant, n2~3; Medium sensitive plant, n4~5; Sensitive plant, NT; Total plant

Seed production of ‘Wonkyo20042ho’ using small net house in 3 years (’12, ’14, and ’15).

Cultivar Location Year Seeds per plant (No.)
Wonkyo20042ho Suwon 2012 1586.1±723
2014 227.6±62.5
Wanju 2015 355.4±46.1

Values are the mean ±SE (n=3) for each year.

Table 1. Collection list of domestic commercial cultivars of Kimchi cabbagez) up to 2007.

Kimchi cabbage is the name of a subgroup of Chinese cabbage which is used as a ingredient of Korean traditional cuisine, Kimchi (codex, 2012).

Table 2. Result of screening for clubroot inoculation using commercial cultivar of Kimchi cabbage.

The number was represented equal cultivar in table 1. Some cultivar was not germinated.

CR test was performed using a single spore isolate with race 4.

Resistance; lv. 1, medium sensitive; lv. 2~3 , sensitive; lv. 4~5 (in Fig. S1)

Table 3. Morphological characteristics of ‘Wonkyo20042ho’.

Color chart: 0; DIC 2087, 1; DIC 2088, 2; DIC 2089, 3; DIC 2540, 4; DIC 167, 5; DIC 166, 6; DIC 86, 7; DIC 85, 8; DIC 84

Color chart: 0; DIC 361, 1; DIC 362, 2; DIC 246, 3; DIC 369, 4; DIC 2324, 5; DIC 2336, 6; DIC 2325, 7; DIC 2356, 8; DIC 2355

Table 4. Variable characteristics of ‘Wonkyo20042ho’.

Values are the mean ±SE (n=3, 3 years are 2012, 2013, 2014).

Table 5. Disease index(DI) z) of ‘Wonkyo20042ho’.

Disease Index (DI) = {(1n1 + 2n2 + 3n3 + 4n4 + 5n5) x 100}/ (4 x NT) n1; Resistance plant, n2~3; Medium sensitive plant, n4~5; Sensitive plant, NT; Total plant

Table 6. Seed production of ‘Wonkyo20042ho’ using small net house in 3 years (’12, ’14, and ’15).

Values are the mean ±SE (n=3) for each year.

Table 1.

Table 2.

Table 3.

Table 4.

Table 5.

Table 6.

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