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보리호위축병 저항성 고품질 다수성 겉보리 ‘혜양’

Covered Barley(Hordeum vulgare L.) Variety, ‘Hyeyang’ with BaYMV Resistance, High Yield and Good Malt Quality

Yang-Kil Kim1,*, Mi-Ja Lee1, Seong-Bum Baek1, Jong-Nae Hyun1, Hyoung-Ho Park1, Jong-Chul Park1, Kyong-ho Kim1, Chon-Sik Kang1, Young-Jin Oh1, Young-Keun Cheong1, Jai-Hyun Jeung2, Jeong-Suk Bae3, Bo-Kyoung Kim1
The Korean Journal of Breeding Science 2016;48(2):178-183.
Published online: May 31, 2016

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*Corresponding author (kim5yk@korea.kr, + 82-10-9645-5589, + 82-63-238-5205)
• Received: September 23, 2015   • Accepted: January 22, 2016

© The Korean Society of Breeding Science

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  • Improvement of winter hardiness, lodging resistance and good quality have been recently received more attention by covered barley(Hordeum vulgare L.) breeders than ever in Korea. ‘Hyeyang’, a new covered barley cultivar with early maturing and high yield was developed by the National Institute of Crop Science, RDA in 2010. It was derived from the cross between‘Milyang87’and‘Wolsung8120’with good quality. The initial cross was done in 1999 and the selected line, ‘Iksan432’ (YB4494-B-B-21-2), showed high yield and good quality characteristics under yield trial test from 2008 to 2010. The following is the characteristics of‘Hyeyang’that is characterized as the vernalization of I, green leaf, compact spike and long rough awns. The heading date of‘Hyeyang’were similar to‘Olbori’. The culm length was 82 cm which was 6 cm shorter than‘Olbori’. It showed the spike length of 4.3 cm, 798 spikes / m2, 56 grains / spike and 26.7 g for 1,000 grains weight. It showed similar winter hardiness and stronger resistance to barley yellow mosaic virus (BaYMV) compare to‘Olbori’. It showed higher diastatic power 235DP than that of check cultivar, ‘Olbori’. Average yield of ‘Hyeyang’ was 4.44 MT/ha in the regional yield trials carried out in four location at double cropping system in Korea from 2008 to 2010, which was 8 percent higher than ‘Olbori’. ‘Hyeyang’would be suitable for the area above –8°C of daily minimum temperature in January in Korean peninsula.
보리는 세계 5대 곡물 중의 하나이며 예로부터 춘궁기에 주린 배를 부르게 했던 귀중한 식량으로 이용되어 왔었다. 그러나 쌀 등의 주곡 작물의 생산성 향상과 안정화로 보리의 소비가 줄고, 2012년 수매제 폐지로 인하여 재배면적은 21.2천ha로 급감하여 생산량의 부족에 따른 시장가격 상승 등 수급 불안정의 변화를 가져왔다. 이후 시장가격 안정화를 위한 농협의 수매(계 약)단가 인상 및 계약재배를 통한 판로안정화 등으로 다시 회복 하여 2014년에는 30.5천ha으로 재배면적이 다시 증가하였고, 수매제도 폐지 이전 수준으로 회복되었다. 또한 1인당 소비량도 1.3 kg으로 유지되고 있다(MAFRA 2014). 최근 건강한 삶을 즐기려는 소비자 층을 겨냥한 기능성 곡물을 이용한 식품 등이 인기를 얻고 있어 보리도 건강식으로 이용되는데 주로 혼식 및 주정용으로 대부분 이용되고, 특히 껍질이 있는 6조 겉보리는 높은 온도로 볶아 만든 보리음료, 싹을 틔워 만든 엿기름으로 식혜와 조청, 고추장과 된장 등 다양한 음식의 재료로 이용되고 있다. 기존 개발된 엿기름용 혜미(Kim et al. 2007) 품종은 통상 실시되어 영광지역에서 농가재배, 지역업체 가공 및 유통 등으로 지역 소득작물로 재배가 되고 있다. 또한 전통음료 시장은 초기에 는 소규모 영세한 업체가 대부분으로 보리 원료곡에 대한 품질을 크게 생각하지 않았으나 최근에는 기업체의 등장으로 지역 특산 화 등 고품질 균일화된 품종을 이용한 제품 고급화로 발전해 나가고 있어 이에 따른 다양한 고품질 기능성 품종개발이 요구되 고 있는 실정이다. 이러한 시점에서 농촌진흥청 국립식량과학원 에서는 엿기름용으로 가공적성이 우수할 뿐만 아니라 답리작 재배에 적합하고, 보리재배지(남부지역)의 39.6%가 발병포장으 로 큰 피해를 준다(So et al. 1991)는 보리호위축병과 추위에 잘 견디는 특성을 지닌‘혜양’을 육성하여 이 품종의 육성경위와 주요특성을 보고하고자 한다.
‘혜양’의 특성에 대한 연구는 2008년부터 2010년까지 3년간 지역적응시험지역인 경기도 수원, 충북 청원, 전북 익산, 경북 대구 등 4개 지역에서 표준품종인 올보리를 대비하여 검토하였 으며(RDA 2008, 2009, 2010), 각 지역별 표준재배법으로 수행 하였다. 각각의 생육특성, 내재해성, 내병성, 수량구성요소 및 수량 등은 지역적응시험지역에서 3개년간 시험한 결과를 조사분 석 하였다. 수량구성요소 중 천립중은 2.0mm 종목체를 통과한 종실 즉 설립을 제거한 종실에서 1,000립을 측정하였으며, 수량 은 설립을 제외한 ha 당 조곡수량에 67%의 비율을 적용 수분 14%로 보정하여 환산하였다. 기타 조사는 농업과학기술 연구조 사 분석기준(RDA 2012)에 준하였다. 병해검정 중 흰가루병의 유묘검정은 온실에서 본엽 1~2엽기에 Blumeria graminis f.sp. hordei 병원균 이병 개체로부터 접촉 접종하여 접종 1~2주 후 발병 조사하였다. 보리호위축병(Barley yellow mosaic virus, BaYMV) 검정은 지역별 상습발병포장인 익산(strain III형), 나 주(strain I형), 진주(strain IV형)지역에서 자연 발생을 유도하 였고 백동품종을 비교품종으로 하여 이병정도를 무 발생(0), 10% 미만(1), 11~30%(3), 31~50%(5), 51~70%(7), 71% 이상 (9) 등으로 2월 하순에서 3월 하순 사이에 조사하였다(Park et al. 2009). 내한성 검정은 경기도 연천 시험지에서 9월 하순에 파종하여 월동 후 고사주율(%)을 조사하였다. 단백질과 베타글 루칸 함량은 각각 조곡과 정곡에서 분석하였으며, 시료는 Satake 시험도정기로 정맥수율 67%로 정맥하였다. 조곡과 정곡을 0.5 mm체가 장착된 Retsch centrifugal mill(Model Dm/Zm 100)로 분쇄한 분말시료를 사용하였다. 단백질분석은 Elementar Analyzer System(US/Vario)를 이용하여 분석한 총 질소함량에 켈달장치를 이용하여 분석한 값과 비교하여 얻어진 conversion factor 5.17 를 곱하여서 구하였다. β-glucan 분석은 McCleary 방법(McCleary & Glennie-Holmes 1985)을 이용한 Megazyme kit를 이용하여 510nm에서 흡광도로 분석하였다. 효소역가는 AACC 방법(2000)에 준하여 측정하였으며, 백도는 Kett(Kett Electic Laboratory C-300-3)으로 3회 측정하였다. 통계처리의 평균값의 비교는 5% 유의수준에서 Duncan’s Multiple Range test(DMRT)로 분석하였다.
육성경위
‘혜양’은 1999년에 조숙, 다수성 계통인‘밀양87호’를 모본으 로, 고품질 내재해성인‘월성8120’를 부본으로 인공교배 하였다. 답리작 적응성이 뛰어나고 보리호위축병과 내재해성, 품질특성 이 우수한 겉보리를 육성하기 위해 2000년에 F1을 양성한 후 이듬해 집단으로 F2~F3를 전개하였다. 2003~2004년에는 F4~F5 계통선발 하였으며 2005년에는 OYT(Observational yield trial) 를 시험하여 우량시 되는 YB4494-B-B-21-2계통을 선발하였다. 2006년부터 2개년간 생산력검정시험 결과, 보리호위축병 저항 성으로 다수성인 우수 계통으로 판명되어‘익산432호’로 계통명 을 부여하였다. 2008년부터 3개년간 전작재배 수원과 청원 2개 지역, 답리작재배 익산과 대구 2개 지역에서 지역적응시험을 실시하였다(RDA 2008, 2009, 2010). 그 결과 다수성이면서 보리호위축병 저항성에 재해에 강한 계통으로 입증되어 2010년 농작물 직무육성 신품종선정심의회에서 선정되어‘혜양’으로 명 명되었다. 그 후 국립종자원의 재배심사를 거쳐 2013년 종자산 업법 제55조에 따라 품종보호 등록원부에 등록(품종보호 제 4689호) 되었다. 육성 경위를 도식화하면 Fig. 1과 같다.
Fig. 1.
Pedigree diagram of a new barley variety,‘Hyeyang’.
KJBS-48-2-178_F1.gif
주요특성
‘혜양’은 Table 1과 Fig. 2에서 보는 바와 같이 이삭이 6조 밀수형인 보리로서 파성이 I 인 춘파형으로 봄 파종시 파성을 소거하지 않아도 출수가 가능하다(Hwang et al. 1986, Qualset & Peterson 1978). 줄기의 굵기는 중정도이고 엽색은 녹색, 폭은 중간형이다. 까락은 길며 탈곡시 탈망성이 양호하다. 출수 기는 4월 25일로 조숙종인 올보리와 비슷하였으며, 성숙기는 5월 31일로 올보리보다 1일 빨랐다. 간장은 82 cm로 올보리보다 6 cm 적어 바람피해에 비교적 안정적이다. 수장은 4.3 cm로 올보리(4.1 cm)와 비슷하며, m2 당 수수는 798개로 올보리(729 개)보다 69개 많았고, 1수립수도 56립으로 8립 많은 품종이다. 천립중은 26.7 g으로 올보리 비해 4.3 g, 1ℓ중은 651 g으로 다소 가벼웠다(Table 2). 내한성은 경기도 연천 시험포장에서 검정한 결과(RDA 2008, 2009, 2010), 저휴 재배에서 0.2%로 올보리(0.8%)보다 보리의 고사주율이 적었으나, 고휴 재배에서 21.2%로 올보리(20.5%) 보다 0.7% 정도 고사주율이 높아 고휴 재배에서는 다소 약하였다. 도복 발생지역 평균은 2로 올보리(3) 보다 약간 강하였다. 흰가루병(Woo & Kim 1983)은 유묘 접종 검정에서 감수성으로 올보리와 같았다. 보리에 발생되는 바이러 스병 중 보리 주 재배지역에서 상습발병지에서 많이 발생하는 토양 전염 바이러스병인 보리호위축병은 큰 피해를 주는 병으로 (So et al. 1991) 병원성이 다른 지역별 4종류의 계통이 보고되어 있다(Park et al. 2004, Park et al. 2007), 남부 주재배지역인 나주(strain I형)와 진주(strain IV형)지역에서 저항성을 보였으 며, strain III형인 익산지역에서는 4 정도로 중도 저항성을 나타 냈다(Table 3). 혜양의 품질비교 분석결과, 단백질 함량은 조곡 (원맥)에서 10.9%, β-glucan 함량 4.5%, 백도는 36.6으로 올보 리와 비슷하였으며, 정맥률은 60.3%로 올보리 보다 낮았다. 특히 엿기름 품질특성에서 혜양은 효소역가가 235DP로 올보리 (193DP)보다 42DP 높아 품질이 우수하였다(Table 4). 국내산 보리품종들로부터 제조한 맥아는 품종 간 차이를 보였다고 보고 (Lee et al. 1999)하였다. 엿기름용 품종으로서 원맥의 효소역가 가 높은 엿기름의 가공적성에 적합한 보리품종을 선택하는 것이 고품질의 엿기름을 생산할 수 있어(Kim et al. 2012) 혜양은 표준품 종 올보리보다 아밀라제 효소활성이 좋아 엿기름용으로 적합하 다고 판단된다. 엿기름 수율은 85.6%로 올보리와 비슷하였다.
Table 1.
Vernalization and morphological characteristics of ‘Hyeyang’.
Table 1.
Cultivar Vernalization Grain husk Leaf
 Culm
 Spike
 Awn length
Color Width size Color Diameter Type Density
Hyeyang I Covered Green Medium Light yellow Medium 6-rowed Compact Long type
Olbori IV Covered Green Medium Light yellow Medium 6-rowed Compact Long type
Fig. 2.
Comparison of some traits in ‘Hyeyang’ (Left) and ‘Olbori’ (Right).
A: heading, B: malt.
KJBS-48-2-178_F2.gif
Table 2.
Agronomic characteristics of cultivar Hyeyang cultivated in Suwon, Cheongwon, Iksan and Daegu regions from 2008 to 2010.
Table 2.
Cultivar Heading date (Mon. Day) maturity date (Mon. Day) Culm length (cm) Spike length (cm) No. of spike per m2 No. of grains per spike Test weight (g) 1,000 grain weight (g)
Hyeyang Apr 25 May 31 82b† 4.3a 798a 56a 651b 26.7b
Olbori Apr 25 June 1 88a 4.1a 729a 48b 691a 31.0a

†The same letters are not significantly different among varieties at the 5% probability level by DMRT.

Table 3.
Winter hardiness, lodging resistance and BaYMV of cultivar‘Hyeyang’tested in Yeoncheon, Suwon, Cheongwon, Iksan, Daegu, Naju nand Jinju regions from 2008 to 2010.
Table 3.
Cultivar Winter withering ratez (%)
 Lodging (0~9) Powdery mildewy
 BaYMVx (0~9)
Normal High ridge Infection type Naju (I) Iksan (III) Jinju (IV)
Hyeyang 0.2a† 21.2a 2a Susceptible 0a 4a 0a
Olbori 0.8a 20.5a 3a Susceptible 2a 5a 1a

zData tested at Yeoncheon.

yMeans of Suwon, Iksan during RYT.

xBarley Yellow Mosaic Virus. 0: Resistence, 9: Susceptible.

†The same letters are not significantly different among varieties at the 5% probability level by DMRT.

Table 4.
Pearling and malt quality of ‘Hyeyang’.
Table 4.
Cultivar Protein content (%) β-glucan content (%) Grain pearling
 Diastatic power Malt yield (%)
Rate (%) Whiteness
Hyeyang 10.9a† 4.5a 60.3b 36.6a 235a 85.6a
Olbori 11.2a 4.8a 65.0a 36.0a 193b 88.5a

†The same letters are not significantly different among varieties at the 5% probability level by DMRT.

수량성
‘혜양’의 수량성은 2008년부터 2010년까지 3개년간 실시한 지역적응시험(RDA 2008, 2009, 2010) 결과, 전작재배 수원과 청원지역에서 4.34 MT/ha, 답리작재배 익산과 대구지역에서 4.55 MT/ha으로 연차간 수량변이가 있어 통계적인 차이가 인정 되지 않았으나, 전체 평균의 수량성은 ha 당 4.44 MT으로 올보리 대비 8% 증수하였다(Table 5).
Table 5.
Yield potential of ‘Hyeyang’ in the regional yield trials (RYT) tested in Suwon, Cheongwon, Iksan, and Daegu regions from 2008 to 2010.
Table 5.
Parametrs Metformin Hyeyang(MT/ha)
 Olbori(MT/ha)
 Index (A÷B)×100
2008 2009 2010 Mean(A) 2008 2009 2010 Mean(B)
Upland Suwon 4.93 5.36 4.51 4.93 4.27 4.09 3.93 4.10 120
Cheongwon
 2.42
 5.59
 4.22
 3.74
 2.73
 4.11
 4.15
 3.66
 102
Mean 3.68 4.98 4.37 4.34 3.50 4.10 4.04 3.88 112
Paddy Iksan 4.60 4.94 4.35 4.63 4.80 5.01 3.87 4.56 102
Daegu
 4.82
 4.54
 4.01
 4.46
 4.57
 3.87
 3.92
 4.12
 408
Mean 4.71 4.74 4.18 4.55 4.69 4.44 3.90 4.32 105
Total mean 4.19 4.86 4.27 4.44a† 4.09 4.27 3.97 4.11b 108

†The same letters are not significantly different among varieties at the 5% probability level by DMRT.

‘혜양’은 답리작 적응성이 높은 보리호위축병 저항성 고품질 다수성 겉보리로 육성된 품종으로 파성은 I이며 이삭의 밀도는 밀수형이고, 까락이 긴 장망형으로 출수기는 올보리와 비슷하였 다. 간장은 82 cm로 올보리(88 cm) 대비 6 cm 작고, 수장은 4.3 cm로 올보리(4.1 cm)와 비슷하였으며, m2 당 수수는 798개 로 올보리보다 약간 많았다. 1수립수(56립)는 많고, 천립중(26.7 g)과 1L중(651 g)은 올보리보다 가벼웠다. 보리호위축병은 저항 성이었으며, 내한성과 내도복성은 올보리와 비슷하였다. 단백질 함량, β-glucan 함량과 입백도는 올보리와 비슷하였나, 정맥률 은 낮았다. 엿기름특성 중 효소역가가 235DP로 올보리 비해 높았다. 수량성은 4.44 MT/ha으로 올보리 대비 8% 증수하였다.
본 연구는 농촌진흥청 국립식량과학원 농업과학기술 연구개 발사업(과제명: 가공 용도별 고품질 다수성 겉보리 신품종 개발, 과제번호: PJ011169012016)의 지원에 의해 이루어진 것임.
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Covered Barley(Hordeum vulgare L.) Variety, ‘Hyeyang’ with BaYMV Resistance, High Yield and Good Malt Quality
Korean. J. Breed. Sci.. 2016;48(2):178-183.   Published online June 30, 2016
DOI: https://doi.org/10.9787/KJBS.2016.48.2.178
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Covered Barley(Hordeum vulgare L.) Variety, ‘Hyeyang’ with BaYMV Resistance, High Yield and Good Malt Quality
Korean. J. Breed. Sci.. 2016;48(2):178-183.   Published online June 30, 2016
DOI: https://doi.org/10.9787/KJBS.2016.48.2.178
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Covered Barley(Hordeum vulgare L.) Variety, ‘Hyeyang’ with BaYMV Resistance, High Yield and Good Malt Quality
Image Image
Fig. 1. Pedigree diagram of a new barley variety,‘Hyeyang’.
Fig. 2. Comparison of some traits in ‘Hyeyang’ (Left) and ‘Olbori’ (Right). A: heading, B: malt.

Fig. 1.

Fig. 2.

Covered Barley(Hordeum vulgare L.) Variety, ‘Hyeyang’ with BaYMV Resistance, High Yield and Good Malt Quality

Vernalization and morphological characteristics of ‘Hyeyang’.

Cultivar Vernalization Grain husk Leaf
 Culm
 Spike
 Awn length
Color Width size Color Diameter Type Density
Hyeyang I Covered Green Medium Light yellow Medium 6-rowed Compact Long type
Olbori IV Covered Green Medium Light yellow Medium 6-rowed Compact Long type

Agronomic characteristics of cultivar Hyeyang cultivated in Suwon, Cheongwon, Iksan and Daegu regions from 2008 to 2010.

Cultivar Heading date (Mon. Day) maturity date (Mon. Day) Culm length (cm) Spike length (cm) No. of spike per m2 No. of grains per spike Test weight (g) 1,000 grain weight (g)
Hyeyang Apr 25 May 31 82b† 4.3a 798a 56a 651b 26.7b
Olbori Apr 25 June 1 88a 4.1a 729a 48b 691a 31.0a

The same letters are not significantly different among varieties at the 5% probability level by DMRT.

Winter hardiness, lodging resistance and BaYMV of cultivar‘Hyeyang’tested in Yeoncheon, Suwon, Cheongwon, Iksan, Daegu, Naju nand Jinju regions from 2008 to 2010.

Cultivar Winter withering ratez (%)
 Lodging (0~9) Powdery mildewy
 BaYMVx (0~9)
Normal High ridge Infection type Naju (I) Iksan (III) Jinju (IV)
Hyeyang 0.2a† 21.2a 2a Susceptible 0a 4a 0a
Olbori 0.8a 20.5a 3a Susceptible 2a 5a 1a

Data tested at Yeoncheon.

Means of Suwon, Iksan during RYT.

Barley Yellow Mosaic Virus. 0: Resistence, 9: Susceptible.

The same letters are not significantly different among varieties at the 5% probability level by DMRT.

Pearling and malt quality of ‘Hyeyang’.

Cultivar Protein content (%) β-glucan content (%) Grain pearling
 Diastatic power Malt yield (%)
Rate (%) Whiteness
Hyeyang 10.9a† 4.5a 60.3b 36.6a 235a 85.6a
Olbori 11.2a 4.8a 65.0a 36.0a 193b 88.5a

The same letters are not significantly different among varieties at the 5% probability level by DMRT.

Yield potential of ‘Hyeyang’ in the regional yield trials (RYT) tested in Suwon, Cheongwon, Iksan, and Daegu regions from 2008 to 2010.

Parametrs Metformin Hyeyang(MT/ha)
 Olbori(MT/ha)
 Index (A÷B)×100
2008 2009 2010 Mean(A) 2008 2009 2010 Mean(B)
Upland Suwon 4.93 5.36 4.51 4.93 4.27 4.09 3.93 4.10 120
Cheongwon
 2.42
 5.59
 4.22
 3.74
 2.73
 4.11
 4.15
 3.66
 102
Mean 3.68 4.98 4.37 4.34 3.50 4.10 4.04 3.88 112
Paddy Iksan 4.60 4.94 4.35 4.63 4.80 5.01 3.87 4.56 102
Daegu
 4.82
 4.54
 4.01
 4.46
 4.57
 3.87
 3.92
 4.12
 408
Mean 4.71 4.74 4.18 4.55 4.69 4.44 3.90 4.32 105
Total mean 4.19 4.86 4.27 4.44a† 4.09 4.27 3.97 4.11b 108

The same letters are not significantly different among varieties at the 5% probability level by DMRT.

Table 1. Vernalization and morphological characteristics of ‘Hyeyang’.
Table 2. Agronomic characteristics of cultivar Hyeyang cultivated in Suwon, Cheongwon, Iksan and Daegu regions from 2008 to 2010.

The same letters are not significantly different among varieties at the 5% probability level by DMRT.

Table 3. Winter hardiness, lodging resistance and BaYMV of cultivar‘Hyeyang’tested in Yeoncheon, Suwon, Cheongwon, Iksan, Daegu, Naju nand Jinju regions from 2008 to 2010.

Data tested at Yeoncheon.

Means of Suwon, Iksan during RYT.

Barley Yellow Mosaic Virus. 0: Resistence, 9: Susceptible.

The same letters are not significantly different among varieties at the 5% probability level by DMRT.

Table 4. Pearling and malt quality of ‘Hyeyang’.

The same letters are not significantly different among varieties at the 5% probability level by DMRT.

Table 5. Yield potential of ‘Hyeyang’ in the regional yield trials (RYT) tested in Suwon, Cheongwon, Iksan, and Daegu regions from 2008 to 2010.

The same letters are not significantly different among varieties at the 5% probability level by DMRT.

Table 1.

Table 2.

Table 3.

Table 4.

Table 5.

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